Before the arrival of the Aborigines, the Canary Islands were inhabited by a variety of endemic animals, such as extinct giant lizards (Gallotia goliath), giant tortoises (Centrochelys burchardi and C. vulcanica), and Tenerife and Gran Canaria giant rats (Canariomys bravoi and C. tamarani), among others. Extinct birds known only from Pleistocene and Holocene age bones include the Canary Islands quail (Coturnix gomerae), Dune shearwater (Puffinus holeae), Lava shearwater (P. olsoni), Trias greenfinch (Chloris triasi), Slender-billed greenfinch (C. aurelioi) and the Long-legged bunting (Emberiza alcoveri). In addition are some unidentified bird eggshell fragments from the Miocene of Lanzarote, which have been suggested to be from an extinct ratite species.